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In 1937, a Chinese martial artist returns to Shanghai to find his teacher dead and his school harassed by the Japanese

An ornithologist investigates reports of a monstrous new species of bird just as a teenage girl is gifted an amulet found on mysterious atoll As the creatures begin to attack, an ancient guardian with a bond to the girl emerges

Gameras ties to humanity have been severed as the Gyaos re emerge Nagamine and Asagi investigate as an orphaned girl named Ayana discovers a new creature Named Iris, Ayana seeks revenge on Gamera for the death of her family